Holtzbrinck Publishing Group is a family-owned German holding company with interests in research publishing (Springer Nature), educational content (Macmillan Learning), consumer books (Macmillan Publishers) and technology (Digital Science, HV Holtzbrinck Ventures). Subsidiaries include ZEIT Group, Nature Publishing Group, Palgrave, Scientific American, St. Martin’s Press, Farrar Straus and Giroux, Henry Holt, S. Fischer Verlage, Rowohlt Verlag, Holtzbrinck Ventures and other fund activities (GSV, Founders Factory, Rethink Education). It has ownership and stakes in start-ups under Digital Science (Dimensions, Overleaf, figshare, Altmetric, Gigantum, überResearch), Macmillan Learning (iClicker, intellus Learning, Achieve), Holtzbrinck Digital (Amboss, pflege.de, lecturio) and the ZEIT Group (Zeit Online, ze.tt).